About the Authors

Paul D Williams

Affiliation Department of Chemistry, University of Michigan, Ann Arbor, Michigan, United States of America

David D Pollock

Affiliation Department of Biological Sciences, Biological Computation and Visualization Center, Louisiana State University, Baton Rouge, Louisiana, United States of America

Benjamin P Blackburne

Affiliation Division of Mathematical Biology, National Institute of Medical Research, Mill Hill, London, United Kingdom

Richard A Goldstein

To whom correspondence should be addressed. E-mail: richard.goldstein@nimr.mrc.ac.uk

Affiliation Division of Mathematical Biology, National Institute of Medical Research, Mill Hill, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

DDP and RAG conceived and designed the experiments. PDW and RAG performed the experiments. PDW, BPB, and RAG analysed the data. BPB and RAG contributed reagents/materials/analysis tools. PDW, DDP, and RAG wrote the paper.