About the Authors

Perttu S Niemelä

To whom correspondence should be addressed. E-mail: psn@fyslab.hut.fi

Affiliations Laboratory of Physics, Helsinki University of Technology, Helsinki, Finland , Helsinki Institute of Physics, Helsinki University of Technology, Helsinki, Finland

Samuli Ollila

Affiliations Laboratory of Physics, Helsinki University of Technology, Helsinki, Finland , Helsinki Institute of Physics, Helsinki University of Technology, Helsinki, Finland

Marja T Hyvönen

Affiliations Laboratory of Physics, Helsinki University of Technology, Helsinki, Finland , Helsinki Institute of Physics, Helsinki University of Technology, Helsinki, Finland , Wihuri Research Institute, Helsinki, Finland

Mikko Karttunen

Affiliation Department of Applied Mathematics, The University of Western Ontario, London, Ontario, Canada

Ilpo Vattulainen

Affiliations Laboratory of Physics, Helsinki University of Technology, Helsinki, Finland , Memphys Center for Biomembrane Physics, University of Southern Denmark, Odense, Denmark , Physics Department, University of Southern Denmark, Odense, Denmark , Institute of Physics, Tampere University of Technology, Tampere, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

PSN, MTH, MK, and IV conceived and designed the experiments. PSN performed the experiments and analyzed the data, except for the lateral pressure profiles computed by SO. PSN wrote the paper together with the other authors.