About the Authors

Sukhbinder Kumar

Affiliation Auditory Group, Medical School, University of Newcastle, Newcastle upon Tyne, United Kingdom

Klaas E Stephan

Affiliation Wellcome Trust Centre for Imaging Neuroscience, Institute of Neurology, University College London, London, United Kingdom

Jason D Warren

Affiliations Wellcome Trust Centre for Imaging Neuroscience, Institute of Neurology, University College London, London, United Kingdom , Dementia Research Centre, Institute of Neurology, University College London, London, United Kingdom

Karl J Friston

Affiliation Wellcome Trust Centre for Imaging Neuroscience, Institute of Neurology, University College London, London, United Kingdom

Timothy D Griffiths

To whom correspondence should be addressed. E-mail: t.d.griffiths@ncl.ac.uk

Affiliations Auditory Group, Medical School, University of Newcastle, Newcastle upon Tyne, United Kingdom , Wellcome Trust Centre for Imaging Neuroscience, Institute of Neurology, University College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

TDG conceived and designed the experiments. SK, KES, JDW, and TDG performed the experiments and analyzed the data. All authors contributed to writing the paper.