About the Authors

Andreas Handel

To whom correspondence should be addressed. E-mail: ahandel@emory.edu

Affiliation Department of Biology, Emory University, Atlanta, Georgia, United States of America

Ira M Longini Jr.

Affiliations Program of Biostatistics and Biomathematics, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America , Department of Biostatistics and School of Public Health and Community Medicine, University of Washington, Seattle, Washington, United States of America

Rustom Antia

Affiliation Department of Biology, Emory University, Atlanta, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

All authors conceived and designed the experiments and wrote the paper. AH performed the experiments and analyzed the data.