About the Authors

Greg J. Stephens

gstephen@princeton.edu (GJS); wsryu@princeton.edu (WSR)

Affiliations Lewis–Sigler Institute for Integrative Genomics, Princeton, New Jersey, United States of America, Joseph Henry Laboratories of Physics, Princeton, New Jersey, United States of America, Center for the Study of Brain, Mind and Behavior, Princeton University, Princeton, New Jersey, United States of America

Bethany Johnson-Kerner

Affiliation Lewis–Sigler Institute for Integrative Genomics, Princeton, New Jersey, United States of America

William Bialek

Affiliations Lewis–Sigler Institute for Integrative Genomics, Princeton, New Jersey, United States of America, Joseph Henry Laboratories of Physics, Princeton, New Jersey, United States of America

William S. Ryu

gstephen@princeton.edu (GJS); wsryu@princeton.edu (WSR)

Affiliation Lewis–Sigler Institute for Integrative Genomics, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WR. Performed the experiments: BJ-K. Wrote the paper: GS WB WR. Analyzed the image data and developed the associated theory: GS WB WR. Analyzed the worm steering data: WR BJ-K.