About the Authors

Yasuko Sugase-Miyamoto

Affiliations Laboratory of Neuropsychology, National Institute of Mental Health, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America, JSPS Overseas Research Fellow, Japan Society for the Promotion of Science, Tokyo, Japan, Neuroscience Research Institute, National Institute of Advanced Industrial Science and Technology, Tsukuba, Japan

Zheng Liu

Affiliation Laboratory of Neuropsychology, National Institute of Mental Health, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Matthew C. Wiener

Affiliation RY84-202, Applied Computer Science and Mathematics Department, Merck Research Laboratories, Rahway, New Jersey, United States of America

Lance M. Optican

Affiliation Laboratory of Sensorimotor Research, Intramural Research Program, National Eye Institute, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Barry J. Richmond

bjr@ln.nimh.nih.gov

Affiliation Laboratory of Neuropsychology, National Institute of Mental Health, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZL BR. Performed the experiments: ZL BR. Analyzed the data: YS ZL MW LO BR. Contributed reagents/materials/analysis tools: YS ZL MW LO BR. Wrote the paper: YS MW LO BR.