About the Authors

Adam B. Barrett

abarrett@inf.ed.ac.uk

Affiliation Institute for Adaptive and Neural Computation, University of Edinburgh, Edinburgh, United Kingdom

Guy O. Billings

Affiliation Institute for Adaptive and Neural Computation, University of Edinburgh, Edinburgh, United Kingdom

Richard G. M. Morris

Affiliation Centre for Cognitive and Neural Systems, University of Edinburgh, Edinburgh, United Kingdom

Mark C. W. van Rossum

Affiliation Institute for Adaptive and Neural Computation, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MCWvR. Performed the experiments: ABB. Analyzed the data: ABB GOB RGMM MCWvR. Wrote the paper: ABB. Came up with the model: ABB. Solved the mathematics: ABB. Helped with designing the model: GOB RGMM MCWvR. Offered comments on draft versions of the paper: GOB. Offered comments on, and helped edit a draft version of the paper: RGMM. Edited the paper: MCWvR.