About the Authors

Nicholas J. Savill

nick.savill@ed.ac.uk

Affiliation Centre for Infectious Diseases, Institute of Immunology and Infection Research, University of Edinburgh, Edinburgh, United Kingdom

William Chadwick

Affiliations Centre for Infectious Diseases, Institute of Immunology and Infection Research, University of Edinburgh, Edinburgh, United Kingdom, Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom

Sarah E. Reece

Affiliations Centre for Infectious Diseases, Institute of Immunology and Infection Research, University of Edinburgh, Edinburgh, United Kingdom, Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NJS WC SER. Performed the experiments: WC SER. Analyzed the data: NJS SER. Wrote the paper: NJS WC SER.