About the Authors

Zhang Zhang

Affiliation Department of Ecology and Evolutionary Biology, Yale University, New Haven, Connecticut, United States of America

Jeffrey P. Townsend

Jeffrey.Townsend@Yale.edu

Affiliation Department of Ecology and Evolutionary Biology, Yale University, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Co-designed and programmed this new method, carried out computer simulations, generated sequence datasets, analyzed the data and wrote the manuscript: ZZ. Designed this new method, supervised the research, and revised the manuscript: JPT.