About the Authors

T. Déirdre Hollingsworth

Contributed equally to this work with: T. Déirdre Hollingsworth, Don Klinkenberg

d.hollingsworth@imperial.ac.uk

Affiliation MRC Centre for Outbreak Control and Analysis, Department of Infectious Disease Epidemiology, Imperial College London, London, United Kingdom

Don Klinkenberg

Contributed equally to this work with: T. Déirdre Hollingsworth, Don Klinkenberg

Affiliation Theoretical Epidemiology, Department of Farm Animal Health, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Hans Heesterbeek

Affiliation Theoretical Epidemiology, Department of Farm Animal Health, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Roy M. Anderson

Affiliation MRC Centre for Outbreak Control and Analysis, Department of Infectious Disease Epidemiology, Imperial College London, London, United Kingdom

Competing Interests

Roy M. Anderson is a non-executive member of the board of GlaxoSmithKline (GSK).

Author Contributions

Conceived and designed the experiments: TDH DK HH RMA. Performed the experiments: TDH DK. Analyzed the data: TDH DK. Wrote the paper: TDH DK HH RMA.