About the Authors

Marcel Salathé

salathe@psu.edu

Affiliation Center for Infectious Disease Dynamics, Department of Biology, Penn State University, University Park, Pennsylvania, United States of America

Shashank Khandelwal

Affiliation Center for Infectious Disease Dynamics, Department of Biology, Penn State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MS. Performed the experiments: MS SK. Analyzed the data: MS SK. Contributed reagents/materials/analysis tools: MS SK. Wrote the paper: MS SK.