About the Authors

Stuart Aitken

s.aitken@ed.ac.uk

Affiliation Centre for Systems Biology, University of Edinburgh, Edinburgh, United Kingdom

Ross D. Alexander

Affiliations Centre for Systems Biology, University of Edinburgh, Edinburgh, United Kingdom, Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Jean D. Beggs

Affiliations Centre for Systems Biology, University of Edinburgh, Edinburgh, United Kingdom, Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RDA JDB. Performed the experiments: RDA. Analyzed the data: SA RDA. Contributed reagents/materials/analysis tools: RDA. Wrote the paper: SA JDB. Performed all modelling, simulation and computational analysis: SA.