About the Authors

Ashok Litwin-Kumar

alk@cmu.edu (ALK); bdoiron@pitt.edu (BD)

Affiliations Program for Neural Computation, Carnegie Mellon University and University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America

Anne-Marie M. Oswald

Affiliations Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America, Department of Biology, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Nathaniel N. Urban

Affiliations Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America, Department of Biology, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Brent Doiron

alk@cmu.edu (ALK); bdoiron@pitt.edu (BD)

Affiliations Center for the Neural Basis of Cognition, Pittsburgh, Pennsylvania, United States of America, Department of Mathematics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ALK AMO NNU BD. Performed the experiments: ALK AMO. Analyzed the data: ALK AMO BD. Wrote the paper: ALK BD.