About the Authors

Renske M. A. Vroomans

Affiliation Theoretical Biology, Utrecht University, Utrecht, The Netherlands

Athanasius F. M. Marée

Affiliation Computational and Systems Biology, John Innes Centre, Norwich Research Park, Norwich, United Kingdom

Rob J. de Boer

Affiliation Theoretical Biology, Utrecht University, Utrecht, The Netherlands

Joost B. Beltman

J.B.Beltman@uu.nl

Affiliations Theoretical Biology, Utrecht University, Utrecht, The Netherlands, Division of Immunology, The Netherlands Cancer Institute, Amsterdam, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RMAV RJB JBB. Performed the experiments: RMAV. Analyzed the data: RMAV AFMM RJB JBB . Contributed reagents/materials/analysis tools: AFMM JBB. Wrote the paper: RMAV AFMM RJB JBB. Designed the environment for the Cellular Potts Model simulations: AFMM.