About the Authors

Yan Zhang

Affiliation Department of Computational Medicine and Bioinformatics, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Hye Kyong Kweon

Affiliation Department of Biological Chemistry, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Christian Shively

Affiliation Department of Molecular, Cellular, and Developmental Biology, University of Michigan, Ann Arbor, Michigan, United States of America

Anuj Kumar

Affiliation Department of Molecular, Cellular, and Developmental Biology, University of Michigan, Ann Arbor, Michigan, United States of America

Philip C. Andrews

andrewsp@umich.edu

Affiliations Department of Computational Medicine and Bioinformatics, University of Michigan Medical School, Ann Arbor, Michigan, United States of America, Department of Biological Chemistry, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: YZ PCA AK. Conceived and initiated the project: PCA YZ AK HKK. Designed and developed the post-identification pipeline, and performed the analysis: YZ. Performed the mass spectrometry experiments: HKK. Contributed biological samples and performed experimental validation: CS. Discussed interpretation of biological results: YZ HKK CS AK PCA.