About the Authors

William R. Harcombe

Affiliation Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America

Nigel F. Delaney

Current address: Department of Medicine, Massachusetts General Hospital, Boston, Massachusetts, United States of America.

Affiliation Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America

Nicholas Leiby

Affiliations Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America, Systems Biology Program, Harvard University, Cambridge, Massachusetts, United States of America

Niels Klitgord

Current address: Life Sciences Division, Lawrence Berkeley National Laboratory, Berkeley, Massachusetts, United States of America.

Affiliation Bioinformatics Graduate Program, Boston University, Boston, Massachusetts, United States of America

Christopher J. Marx

cmarx@oeb.harvard.edu

Affiliations Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America, Faculty of Arts and Sciences Center for Systems Biology, Harvard University, Cambridge, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WRH CJM. Performed the experiments: WRH. Analyzed the data: WRH NFD NL NK CJM. Contributed reagents/materials/analysis tools: WRH NFD NK. Wrote the paper: WRH NFD NL NK CJM.