About the Authors

Harish Shankaran

Affiliation Computational Biology and Bioinformatics Group, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Yi Zhang

Affiliation Computational Biology and Bioinformatics Group, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Yunbing Tan

Affiliation School of Electrical Engineering and Computer Science, Washington State University, Pullman, Washington, United States of America

Haluk Resat

haluk.resat@wsu.edu

Current address: School of Chemical Engineering and Bioengineering, Washington State University, Pullman, Washington, United States of America.

Affiliation Computational Biology and Bioinformatics Group, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS YZ HR. Performed the experiments: YZ. Analyzed the data: HS YT HR. Wrote the paper: HS HR. Coded the mathematical model: HS YT.