About the Authors

Yuanyuan Xiao

Contributed equally to this work with: Yuanyuan Xiao, Yee H Yang

Affiliation Department of Epidemiology and Biostatistics, Center for Bioinformatics and Molecular Biostatistics, University of California, San Francisco, California, United States of America

Yee H Yang

Contributed equally to this work with: Yuanyuan Xiao, Yee H Yang

Affiliation Department of Medicine, Center for Bioinformatics and Molecular Biostatistics, University of California, San Francisco, California, United States of America

Todd A Burckin

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California, Santa Cruz, California, United States of America

Lily Shiue

Affiliation Department of Molecular, Cell and Developmental Biology, University of California, Santa Cruz, California, United States of America

Grant A Hartzog

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California, Santa Cruz, California, United States of America

Mark R Segal

To whom correspondence should be addressed. E-mail: mark@biostat.ucsf.edu

Affiliation Department of Epidemiology and Biostatistics, Center for Bioinformatics and Molecular Biostatistics, University of California, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

GAH and MRS conceived and designed the experiments. TAB performed the experiments. YX, YHY, and MRS analyzed the data. YHY, LS, and GAH contributed reagents/materials/analysis tools. YX, YHY, TAB, GAH, and MRS wrote the paper.