About the Authors

Vitaly V Ganusov

Contributed equally to this work with: Vitaly V Ganusov, Rustom Antia

¤ Current address: Theoretical Biology, Utrecht University, Utrecht, Netherlands

Affiliation Department of Biology, Emory University, Atlanta, Georgia, United States of America

Sergei S Pilyugin

Affiliation Department of Mathematics, University of Florida, Gainesville Florida, United States of America

Rafi Ahmed

Affiliation Emory Vaccine Center, Emory University, Atlanta, Georgia, United States of America

Rustom Antia

Contributed equally to this work with: Vitaly V Ganusov, Rustom Antia

To whom correspondence should be addressed. E-mail: rantia@emory.edu

Affiliation Department of Biology, Emory University, Atlanta, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

R. Antia and V. V. Ganusov conceived the basic ideas. R. Ahmed helped formulate the problem and consider the implications for experimental work. R. Antia and V. V. Ganusov derived analytical approximations. V. V. Ganusov performed the simulations. V. V. Ganusov and R. Antia wrote the paper. S. S. Pilyugin contributed to the derivations and helped to modify the text.