About the Authors

John W Pepper

To whom correspondence should be addressed. E-mail: jpepper1@email.arizona.edu

Affiliations Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, Arizona, United States of America , The Santa Fe Institute, Santa Fe, New Mexico, United States of America

Kathleen Sprouffske

Affiliations Genomics and Computational Biology Program, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America , Systems Biology Division, The Wistar Institute, Philadelphia, Pennsylvania, United States of America

Carlo C Maley

Affiliation Systems Biology Division, The Wistar Institute, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JWP and CCM conceived and designed the experiments. JWP, KS, and CCM performed the experiments. JWP, KS, and CCM analyzed the data. JWP and CCM wrote the paper.