About the Authors

Robert M Seymour

Affiliations Department of Mathematics, University College London, London, United Kingdom , Centre for Mathematics and Physics in the Life Sciences and Experimental Biology (CoMPLEX), University College London, London, United Kingdom

C. Patrick Doncaster

To whom correspondence should be addressed. E-mail: cpd@soton.ac.uk

Affiliation School of Biological Sciences, University of Southampton, Southampton, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

RMS conceived and designed the model. CPD conceived and designed the simulations. RMS and CPD wrote the paper.