About the Authors

Shobhit Gupta

Contributed equally to this work with: Shobhit Gupta, Jonathan Dennis

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Jonathan Dennis

Contributed equally to this work with: Shobhit Gupta, Jonathan Dennis

Affiliation Department of Molecular Biology, Massachusetts General Hospital, Boston, Massachusetts, United States of America

Robert E. Thurman

Affiliation Division of Medical Genetics, University of Washington, Seattle, Washington, United States of America

Robert Kingston

Affiliation Department of Molecular Biology, Massachusetts General Hospital, Boston, Massachusetts, United States of America

John A. Stamatoyannopoulos

jstam@u.washington.edu; noble@gs.washington.edu

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

William Stafford Noble

jstam@u.washington.edu; noble@gs.washington.edu

Affiliations Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America, Department of Computer Science and Engineering, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SG JD JAS WSN. Performed the experiments: SG JD. Analyzed the data: SG JD RET. Wrote the paper: SG JD RK JAS WSN.