About the Authors

John Archer

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Andrew Rambaut

Affiliation Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom

Bruce E. Taillon

Affiliation 454 Life Sciences, Branford, Connecticut, United States of America

P. Richard Harrigan

Affiliations British Columbia Centre for Excellence in HIV/AIDS, Vancouver, British Columbia, Canada, Faculty of Medicine, University of British Columbia, Vancouver, British Columbia, Canada

Marilyn Lewis

Affiliation Pfizer Global Research and Development, Sandwich, United Kingdom

David L. Robertson

david.robertson@manchester.ac.uk

Affiliation Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ML DLR. Performed the experiments: JA BET PRH. Analyzed the data: JA DLR. Contributed reagents/materials/analysis tools: AR. Wrote the paper: JA ML DLR.