About the Authors

C. David Williams

cdave@uw.edu

Affiliation Department of Physiology and Biophysics, University of Washington, Seattle, Washington, United States of America

Michael Regnier

Affiliations Department of Physiology and Biophysics, University of Washington, Seattle, Washington, United States of America, Department of Bioengineering, University of Washington, Seattle, Washington, United States of America

Thomas L. Daniel

Affiliations Department of Bioengineering, University of Washington, Seattle, Washington, United States of America, Department of Biology, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CDW MR TLD. Performed the experiments: CDW. Analyzed the data: CDW TLD. Contributed reagents/materials/analysis tools: CDW TLD. Wrote the paper: CDW MR TLD.