About the Authors

Sebastian Funk

sf7@princeton.edu

Affiliations Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America, London School of Hygiene & Tropical Medicine, London, United Kingdom

Hiroshi Nishiura

Affiliations School of Public Health, The University of Hong Kong, Hong Kong, People's Republic of China, PRESTO, Japan Science of Technology Agency, Kawaguchi, Saitama, Japan

Hans Heesterbeek

Affiliation Theoretical Epidemiology, Faculty of Veterinary Medicine, University of Utrecht, Utrecht, The Netherlands

W. John Edmunds

Affiliation London School of Hygiene & Tropical Medicine, London, United Kingdom

Francesco Checchi

Affiliation London School of Hygiene & Tropical Medicine, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Developed the methods: SF HN HH. Analyzed the data: SF. Wrote the paper: SF HN HH WJE FC.