About the Authors

Kieran Alden

k.j.alden@bham.ac.uk

Affiliation Centre for Systems Biology, School of Biosciences, University of Birmingham, Birmingham, United Kingdom

Mark Read

Affiliation Department of Electronics, University of York, York, United Kingdom

Jon Timmis

Affiliations Department of Electronics, University of York, York, United Kingdom, Department of Computer Science, University of York, York, United Kingdom

Paul S. Andrews

Affiliation Department of Computer Science, University of York, York, United Kingdom

Henrique Veiga-Fernandes

Affiliation Instituto de Medicina Molecular, Faculdade de Medicina de Lisboa, Lisboa, Portugal

Mark Coles

Affiliation Centre for Immunology and Infection, University of York and Hull York Medical School, York, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KA MR JT PSA MC. Performed the experiments: KA. Analyzed the data: KA MR. Contributed reagents/materials/analysis tools: MR HVF MC. Wrote the paper: KA MR JT PSA HVF MC. Provided biological expertise in the development of the simulator and understanding of the results it generated: HVF MC.