About the Authors

Yuan-Yuan Li

Contributed equally to this work with: Yuan-Yuan Li, Hui Yu

Affiliation Shanghai Center for Bioinformation Technology, Shanghai, People's Republic of China

Hui Yu

Contributed equally to this work with: Yuan-Yuan Li, Hui Yu

Affiliation Shanghai Center for Bioinformation Technology, Shanghai, People's Republic of China

Zong-Ming Guo

Affiliation Shanghai Center for Bioinformation Technology, Shanghai, People's Republic of China

Ting-Qing Guo

Affiliation Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, People's Republic of China

Kang Tu

Affiliations Shanghai Center for Bioinformation Technology, Shanghai, People's Republic of China , Bioinformatics Center, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, People's Republic of China

Yi-Xue Li

To whom correspondence should be addressed. E-mail: yxli@scbit.org

Affiliations Shanghai Center for Bioinformation Technology, Shanghai, People's Republic of China , Bioinformatics Center, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

YYL and YXL conceived and designed the experiments. HY, ZMG, and KT performed the experiments. YYL, HY, and TQG analyzed the data. YXL managed and supervised the project. YYL and HY wrote the paper.