About the Authors

Hideaki Ogasawara

To whom correspondence should be addressed. E-mail: ogahide@atr.jp

Affiliations ATR Computational Neuroscience Laboratories, Seika, Kyoto, Japan , National Institute of Information and Communications Technology, Seika, Kyoto, Japan

Tomokazu Doi

¤ Current address: Systems Biology Department, Osaka Bioscience Institute, Suita, Osaka, Japan

Affiliations ATR Computational Neuroscience Laboratories, Seika, Kyoto, Japan , Graduate School of Information Science, Nara Institute of Science and Technology, Takayama, Ikoma, Nara, Japan

Kenji Doya

Affiliations ATR Computational Neuroscience Laboratories, Seika, Kyoto, Japan , Initial Research Project, Okinawa Institute of Science and Technology, Uruma, Okinawa, Japan

Mitsuo Kawato

Affiliation ATR Computational Neuroscience Laboratories, Seika, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

HO, TD, KD, and MK conceived and designed the experiments. HO performed the experiments. HO and MK analyzed the data. HO and TD contributed reagents/materials/analysis tools. HO and MK wrote the paper.