About the Authors

Philippe Lemey

To whom correspondence should be addressed. E-mail: philippe.lemey@zoo.ox.ac.uk

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Sergei L Kosakovsky Pond

Affiliation Department of Pathology, University of California San Diego, La Jolla, United States of America

Alexei J Drummond

Affiliation Department of Computer Science, University of Auckland, Auckland, New Zealand

Oliver G Pybus

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Beth Shapiro

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Helena Barroso

Affiliations Centro de Patogénese Molecular, Faculdade de Farmácia de Lisboa, Lisbon, Portugal , Instituto Superior de Ciências da Saúde Egas Moniz, Lisbon, Portugal

Nuno Taveira

Affiliations Centro de Patogénese Molecular, Faculdade de Farmácia de Lisboa, Lisbon, Portugal , Instituto Superior de Ciências da Saúde Egas Moniz, Lisbon, Portugal

Andrew Rambaut

Affiliation Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

PL, OGP, and AR designed the analysis strategy. PL analysed the data and wrote the paper. SLKP assisted in HYPHY analyses. AJD and AR provided programming assistance. BS, AJD, and AR assisted in BEAST analyses. NT and HB contributed unpublished HIV-2 transmission chain data.