About the Authors

Gert Jan Boender

Affiliation Animal Sciences Group, Wageningen University and Research Centre, Lelystad, The Netherlands

Thomas J Hagenaars

Affiliation Animal Sciences Group, Wageningen University and Research Centre, Lelystad, The Netherlands

Annemarie Bouma

Affiliation Faculty of Veterinary Medicine, Department of Farm Animal Health, Utrecht University, Utrecht, The Netherlands

Gonnie Nodelijk

Affiliation Animal Sciences Group, Wageningen University and Research Centre, Lelystad, The Netherlands

Armin R. W Elbers

¤ Current address: Department of Virology, Central Institute for Animal Disease Control, Wageningen University and Research Centre, Lelystad, The Netherlands

Affiliation Animal Health Service, Deventer, The Netherlands

Mart C. M de Jong

Affiliation Animal Sciences Group, Wageningen University and Research Centre, Lelystad, The Netherlands

Michiel van Boven

To whom correspondence should be addressed. E-mail: michiel.vanboven@wur.nl

Affiliation Animal Sciences Group, Wageningen University and Research Centre, Lelystad, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MCMdJ, GJB, MvB, and TJH conceived the ideas underlying the analyses. AB, GN, and ARWE collected the data. GJB, TJH, and MvB analyzed the data and wrote the paper. AB, GN, ARWE, and MCMdJ commented on the text.