About the Authors

Harish Shankaran

Affiliation Systems Biology Program, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Haluk Resat

To whom correspondence should be addressed. E-mail: haluk.resat@pnl.gov

Affiliation Systems Biology Program, Pacific Northwest National Laboratory, Richland, Washington, United States of America

H. Steven Wiley

Affiliation Systems Biology Program, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

HS, HR, and HSW conceived and designed the experiments and wrote the paper. HS performed the experiments.