About the Authors

Naoki Masuda

To whom correspondence should be addressed. E-mail: masuda@mist.i.u-tokyo.ac.jp (NM), bdoiron@pitt.edu (BD)

Affiliations Graduate School of Information Science and Technology, the University of Tokyo, Bunkyo, Tokyo, Japan , Amari Research Unit, RIKEN Brain Science Institute, Wako, Saitama, Japan

Brent Doiron

To whom correspondence should be addressed. E-mail: masuda@mist.i.u-tokyo.ac.jp (NM), bdoiron@pitt.edu (BD)

¤ Current address: Department of Mathematics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Affiliation Center for Neural Science, New York University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

NM and BD conceived and designed the experiments. NM and BD performed the experiments. NM and BD wrote the paper.