About the Authors

Clinton H Hansen

Affiliation Department of Physics, Princeton University, Princeton, New Jersey, United States of America

Robert G Endres

¤ Current address: Division of Molecular Biosciences and CISBIC, Imperial College, London, United Kingdom

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Ned S Wingreen

To whom correspondence should be addressed. E-mail: wingreen@princeton.edu

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

CHH, RGE, and NSW analyzed data and the literature, RGE and NSW conceived and designed the model. CHH performed the model calculations. CHH, RGE, and NSW wrote the paper.