About the Authors

Christian L. Althaus

c.l.althaus@uu.nl

Rob J. De Boer

Affiliation Theoretical Biology, Utrecht University, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CA RDB. Performed the experiments: CA. Analyzed the data: CA. Contributed reagents/materials/analysis tools: CA. Wrote the paper: CA.