About the Authors

Zhengdong D. Zhang

Affiliation Department of Molecular Biophysics and Biochemistry, Yale University, New Haven, Connecticut, United States of America

Joel Rozowsky

Affiliation Department of Molecular Biophysics and Biochemistry, Yale University, New Haven, Connecticut, United States of America

Michael Snyder

Affiliation Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America

Joseph Chang

Affiliation Department of Statistics, Yale University, New Haven, Connecticut, United States of America

Mark Gerstein

mark.gerstein@yale.edu

Affiliations Department of Molecular Biophysics and Biochemistry, Yale University, New Haven, Connecticut, United States of America, Interdepartmental Program in Computational Biology and Bioinformatics, Yale University, New Haven, Connecticut, United States of America, Department of Computer Science, Yale University, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZDZ JC. Performed the experiments: ZDZ. Analyzed the data: ZDZ. Contributed reagents/materials/analysis tools: JR MS. Wrote the paper: ZDZ MBG.