About the Authors

Yuki Tsukada

Affiliations Laboratory for Systems Biology, Graduate School of Information Science, Nara Institute of Science and Technology, Nara, Japan, Institute for Bioinformatics Research and Development, Japan Science and Technology Agency, Tokyo, Japan

Kazuhiro Aoki

Affiliation Laboratory of Bioimaging and Cell Signaling, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Takeshi Nakamura

Affiliation Laboratory of Bioimaging and Cell Signaling, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Yuichi Sakumura

Affiliations Laboratory for Systems Biology, Graduate School of Information Science, Nara Institute of Science and Technology, Nara, Japan, Institute for Bioinformatics Research and Development, Japan Science and Technology Agency, Tokyo, Japan

Michiyuki Matsuda

Affiliation Laboratory of Bioimaging and Cell Signaling, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Shin Ishii

ishii@i.kyoto-u.ac.jp

Affiliations Laboratory for Systems Biology, Graduate School of Information Science, Nara Institute of Science and Technology, Nara, Japan, Institute for Bioinformatics Research and Development, Japan Science and Technology Agency, Tokyo, Japan, Integrated Systems Biology Laboratory, Department of Systems Science, Graduate School of Informatics, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YT KA TN YS. Performed the experiments: KA MM. Analyzed the data: YT. Contributed reagents/materials/analysis tools: YT. Wrote the paper: YT KA TN YS MM SI.