About the Authors

Matthew J. Paszek

Affiliations Department of Bioengineering, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Institute for Medicine and Engineering, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Center for Bioengineering and Tissue Regeneration, Department of Surgery, University of California, San Francisco, San Francisco, California, United States of America

David Boettiger

Affiliations Institute for Medicine and Engineering, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Microbiology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Pharmacology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Valerie M. Weaver

Affiliations Center for Bioengineering and Tissue Regeneration, Department of Surgery, University of California, San Francisco, San Francisco, California, United States of America, Department of Anatomy, University of California, San Francisco, San Francisco, California, United States of America, Department of Bioengineering and Therapeutic Sciences, Institute for Regenerative Medicine and UCSF Comprehensive Cancer Center, University of California, San Francisco, San Francisco, California, United States of America

Daniel A. Hammer

hammer@seas.upenn.edu

Affiliations Department of Bioengineering, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Chemical and Biomolecular Engineering, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Institute for Medicine and Engineering, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DAH. Performed the experiments: MJP. Analyzed the data: DB VMW. Wrote the paper: MJP DB VMW. Supervised the work: DAH. Helped write the paper: DAH. Assisted with analysis: DAH.