About the Authors

Jun Cai

Current address: Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China

Affiliation Department of Bioengineering, University of Illinois at Urbana Champaign, Urbana, Illinois, United States of America

Dan Xie

Affiliation Department of Bioengineering, University of Illinois at Urbana Champaign, Urbana, Illinois, United States of America

Zhewen Fan

Affiliations Department of Bioengineering, University of Illinois at Urbana Champaign, Urbana, Illinois, United States of America, Department of Statistics, University of Illinois at Urbana Champaign, Urbana, Illinois, United States of America

Hiram Chipperfield

Affiliation Institute of Medical Biology, Singapore

John Marden

Affiliation Department of Statistics, University of Illinois at Urbana Champaign, Urbana, Illinois, United States of America

Wing H. Wong

Affiliations Department of Statistics, Stanford University, Stanford, California, United States of America, Department of Health Research and Policy, Stanford University, Stanford, California, United States of America

Sheng Zhong

szhong@illinois.edu

Affiliations Department of Bioengineering, University of Illinois at Urbana Champaign, Urbana, Illinois, United States of America, Department of Statistics, University of Illinois at Urbana Champaign, Urbana, Illinois, United States of America, Institute of Genomic Biology, University of Illinois at Urbana Champaign, Urbana, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SZ. Performed the experiments: HC WHW. Analyzed the data: JC DX ZF SZ. Contributed reagents/materials/analysis tools: JC DX ZF JM WHW SZ. Wrote the paper: SZ.