About the Authors

Jonathan E. Kohn

Affiliation Department of Bioengineering, University of California, Berkeley, Berkeley, California, United States of America

Pavel V. Afonine

Affiliation Physical Biosciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Jory Z. Ruscio

Affiliation Department of Bioengineering, University of California, Berkeley, Berkeley, California, United States of America

Paul D. Adams

Affiliations Department of Bioengineering, University of California, Berkeley, Berkeley, California, United States of America, Physical Biosciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Teresa Head-Gordon

TLHead-Gordon@lbl.gov

Affiliations Department of Bioengineering, University of California, Berkeley, Berkeley, California, United States of America, Physical Biosciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Competing Interests

JZR was employed by the Public Library of Science at the time of submission.

Author Contributions

Conceived and designed the experiments: THG. Performed the experiments: JEK PVA JZR. Analyzed the data: JEK PDA THG. Wrote the paper: THG.