About the Authors

Wenyuan Li

Affiliation Molecular and Computational Biology, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America

Chun-Chi Liu

Affiliation Molecular and Computational Biology, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America

Tong Zhang

Affiliation Department of Statistics, Rutgers University, New Brunswick, New Jersey, United States of America

Haifeng Li

Affiliation Molecular and Computational Biology, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America

Michael S. Waterman

Affiliation Molecular and Computational Biology, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America

Xianghong Jasmine Zhou

xjzhou@usc.edu

Affiliation Molecular and Computational Biology, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WL XJZ. Performed the experiments: WL XJZ. Analyzed the data: WL CCL XJZ. Contributed reagents/materials/analysis tools: WL TZ XJZ. Wrote the paper: WL CCL XJZ. Initiated the research: HL MSW.