About the Authors

Helen R. Fryer

helen.fryer@zoo.ox.ac.uk

Affiliation The Institute for Emerging Infections, The Oxford Martin School, Department of Zoology, Oxford University, Oxford, United Kingdom

Angela R. McLean

Affiliation The Institute for Emerging Infections, The Oxford Martin School, Department of Zoology, Oxford University, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: HRF ARM. Conceived and designed the study: ARM HRF. Performed the mathematical modelling and analysis: HRF.