About the Authors

Karl J. Friston

k.friston@fil.ion.ucl.ac.uk

Affiliation The Wellcome Trust Centre for Neuroimaging, University College London, Queen Square, London, United Kingdom

Tamara Shiner

Affiliation The Wellcome Trust Centre for Neuroimaging, University College London, Queen Square, London, United Kingdom

Thomas FitzGerald

Affiliation The Wellcome Trust Centre for Neuroimaging, University College London, Queen Square, London, United Kingdom

Joseph M. Galea

Affiliation Sobell Department of Motor Neuroscience and Movement Disorders, University College London Institute of Neurology, Queen Square, London, United Kingdom

Rick Adams

Affiliation The Wellcome Trust Centre for Neuroimaging, University College London, Queen Square, London, United Kingdom

Harriet Brown

Affiliation The Wellcome Trust Centre for Neuroimaging, University College London, Queen Square, London, United Kingdom

Raymond J. Dolan

Affiliation The Wellcome Trust Centre for Neuroimaging, University College London, Queen Square, London, United Kingdom

Rosalyn Moran

Affiliation The Wellcome Trust Centre for Neuroimaging, University College London, Queen Square, London, United Kingdom

Klaas Enno Stephan

Affiliation The Wellcome Trust Centre for Neuroimaging, University College London, Queen Square, London, United Kingdom

Sven Bestmann

Affiliation Sobell Department of Motor Neuroscience and Movement Disorders, University College London Institute of Neurology, Queen Square, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KF JG SB. Performed the experiments: KF. Wrote the paper: KF TS TF JG RA HB RD RM KS SB.