About the Authors

Caroline O. Buckee

Affiliation Center for Communicable Disease Dynamics, Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America

Mario Recker

mario.recker@zoo.ox.ac.uk

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: COB MR. Analyzed the data: COB. Wrote the paper: COB MR. Designed the models and analysis: COB MR. Performed the statistical analysis of the sequence data: COB. Performed the mathematical models: MR.