About the Authors

Yasuhiro Tsubo

yasuhirotsubo@riken.jp

Affiliation Laboratory for Neural Circuit Theory, RIKEN Brain Science Institute, Wako, Saitama, Japan

Yoshikazu Isomura

Affiliations Laboratory for Neural Circuit Theory, RIKEN Brain Science Institute, Wako, Saitama, Japan, Brain Science Institute, Tamagawa University, Machida, Tokyo, Japan, CREST, JST, 4-1-8 Honcho, Kawaguchi, Saitama, Japan

Tomoki Fukai

Affiliations Laboratory for Neural Circuit Theory, RIKEN Brain Science Institute, Wako, Saitama, Japan, CREST, JST, 4-1-8 Honcho, Kawaguchi, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YT YI TF. Performed the experiments: YI. Analyzed the data: YT. Wrote the paper: YT TF.