About the Authors

Jayanthi Santhanam

jsanthan@staffmail.ed.ac.uk

Affiliation Institute of Immunology and Infection Research, University of Edinburgh, Ashworth Labs, Edinburgh, Scotland

Lars Råberg

Affiliation Department of Biology, Lund University, Lund, Sweden

Andrew F. Read

Affiliation Center for Infectious Disease Dynamics, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Nicholas Jon Savill

Affiliation Institute of Immunology and Infection Research, University of Edinburgh, Ashworth Labs, Edinburgh, Scotland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NJS. Performed the experiments: JS. Analyzed the data: JS NJS. Contributed reagents/materials/analysis tools: LR AFR. Wrote the paper: JS LR AFR NJS.