Isolation-by-Distance and Outbreeding Depression Are Sufficient to Drive Parapatric Speciation in the Absence of Environmental Influences

• Guy A. Hoelzer ,

  * E-mail: hoelzer@unr.edu

  Affiliation Department of Biology, University of Nevada, Reno, Nevada, United States of America

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• Rich Drewes,

  Affiliation Biomedical Engineering Program, University of Nevada, Reno, Nevada, United States of America

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• Jeffrey Meier,

  Affiliation Department of Mathematics and Statistics, University of Nevada, Reno, Nevada, United States of America

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• René Doursat

  Affiliation Institut des Systèmes Complexes, CREA, CNRS, and Ecole Polytechnique, Paris, France

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